| Main Ref. | Takeshita, N., N. Onikura, S. Nagata, S. Matsui and S. Kimura, 1999 |
| Mode | dioecism |
| Fertilization | external |
| Mating type | promiscuity |
| Spawning frequency | |
| Spawning aggregation | Ref. |
| Batch spawner | Ref. |
| Reproductive guild |
guarders clutch tenders |
| Parental Care | paternal |
| Description of life cycle and mating behavior | |
| Search for more references on reproduction | Scirus |