| Main Ref. | Sokolov, L.I. and L.S. Berdicheskii, 1989 |
| Mode | dioecism |
| Fertilization | external |
| Mating type | |
| Spawning frequency | |
| Spawning aggregation | Ref. |
| Batch spawner | Ref. |
| Reproductive guild |
nonguarders open water/substratum egg scatterers |
| Parental Care | none |
| Description of life cycle and mating behavior | Mature adults make upstream spawning migrations beginning as early as June and may extend to September or later (Ref. 28214, 28852, 28855, 28857) and move back downstream after spawning and overwinter in deep parts of the rivers or in estuaries (Ref. 27547). Young hatch in the spring and move downward (Ref. 27547). |
| Search for more references on reproduction | Scirus |