| Main Ref. | Toshitaka, H. and S. Takahashi, 1987 |
| Mode | dioecism |
| Fertilization | external |
| Mating type | monogamy: |
| Spawning frequency | once in a lifetime |
| Spawning aggregation | Ref. |
| Batch spawner | Ref. Nakazawa, T., N. Ishida, M. Kato and N. Yamamura, 2007 |
| Reproductive guild |
guarders nesters |
| Parental Care | paternal |
| Description of life cycle and mating behavior | Migrate to the shore to spawn under stones in spring (Ref. 76915). Males have only two brood cycles, mating with the same female during each breeding cycle (Ref. 76915). |
| Search for more references on reproduction | Scirus |