This species is characterized by the following: usual number of vertebrae, precaudal 14-15, caudal 65-67, total 81-82 (80-83); usual number of fin rays, dorsal 142-145 (140-147), anal 120-125, pectoral 22-23); pyloric caeca usually 3 (rarely 4), 2 (rarely 3) in a single whorl, 1 in 2nd tier; first gill arch with rudimentary rakers 4 (rarely 3) on the upper arm, 8 developed (rarely 7) developed rakers on the lower limb, 0-3 (usually 0) rudimentary rakers anteriorly on the first arch; total rakers 12 (usually 11-14); dorsal-fin origin between neural spines 4-5 (11 specimens) or 5-6 (6 specimens) (Ref. 91765).