| Main Ref. | Karino, K. and A. Nakazono, 1993 |
| Mode | dioecism |
| Fertilization | external |
| Mating type | promiscuity |
| Spawning frequency | one clear seasonal peak per year |
| Spawning aggregation | Ref. |
| Batch spawner | Ref. |
| Reproductive guild |
guarders nesters |
| Parental Care | paternal |
| Description of life cycle and mating behavior | Oviparous, distinct pairing during breeding (Ref. 205). Eggs are demersal and adhere to the substrate (Ref. 205). Males guard and aerate the eggs (Ref. 205). |
| Search for more references on reproduction | Scirus |