| Main Ref. | Kullander, S.O., F. Fang, B. Delling and E. Åhlander, 1999 |
| Mode | dioecism |
| Fertilization | external |
| Mating type | |
| Spawning frequency | |
| Spawning aggregation | Ref. |
| Batch spawner | Ref. |
| Reproductive guild |
nonguarders open water/substratum egg scatterers |
| Parental Care | none |
| Description of life cycle and mating behavior | Mature adults undertake spawning migration to incoming streams where they breed amidst gravel and sandy beds (Ref. 40974). |
| Search for more references on reproduction | Scirus |